Oliver's mTOR Atlas Evidence Platform
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mTOR Academy · Practice

Practice Arena

Eight games built on the Atlas's own pathway model and its own studies. Points reward calibration, not speed — and nothing here rewards simply showing up.

48 nodes in the Academy map · 88 in the Atlas

The pathway is the board you are playing on. Every question you answer colours in a piece of it — and two of its questions are ones the field itself has not closed, so they never fill in.

Your pathway and badges →

Rank 1

Curious

0 XPProgress is kept in this browser
mixed

Daily 5

Two you have seen, two new, one a level up. Three minutes.

Recall

Signal Sprint

Sixty seconds of arrows: does this one activate or inhibit?

Structure

Wire the Pathway

Rebuild a route from loose parts — and get the direct steps right.

Prediction

Perturbation Lab

Set the controls, predict the readout before it is revealed.

Limits of evidence

What It Doesn't Show

One result, four readings. Which one the evidence will not carry?

Reading methods

Paper Autopsy

One real result, four readings. Which one the paper cannot carry?

Traceability

Find the Evidence

A claim from the map. Which study in the Atlas is it standing on?

State of knowledge

Frontier

Established, emerging, contested or open? Say where the field actually is.

How the points work

Points are called XP. Every number you see with a plus sign — +8 XP, +30 XP — is XP added to the total in the bar above; the ranks are XP thresholds. Every question asks how sure you are before the answer appears. Being right while sure is worth the most; being right while unsure is worth less; being wrong while sure is worth nothing at all. There are no points for opening a page, no daily-login bonus and no streak to lose — the only thing that earns anything here is a judgement that could have been wrong.

Open questions never reach full mastery. That is not a bug in the game — the field has not closed them.

Practice questions

This page needs JavaScript for scoring and for saving progress in this browser. The practice questions themselves are below in full, with answers — the page is readable and usable without it.

What kind of molecule is mTOR?
  1. A hormone the body releases when food is plentiful
  2. A protein kinase — an enzyme that attaches phosphate groups to other proteins
  3. A transcription factor that binds DNA at growth genes
  4. A structural protein that holds the lysosome together

Answer: B. mTOR is an enzyme. It changes what other proteins do by phosphorylating them, and everything downstream in this course follows from that one activity.

Which subunit is the one that defines mTORC1 rather than mTORC2?
  1. RICTOR
  2. SIN1
  3. RAPTOR
  4. mLST8

Answer: C. RAPTOR is the mTORC1-defining partner and RICTOR and SIN1 belong to mTORC2. mLST8 is the distractor worth remembering, because it sits in both complexes.

Which form of Rheb is the direct positive input to mTORC1 in the standard model?
  1. Rheb-GDP
  2. Rheb-GTP
  3. Both forms equally
  4. Neither — Rheb acts only on mTORC2

Answer: B. Rheb is a switch protein whose two nucleotide states behave differently, and it is the GTP-bound state that activates the kinase. Everything the TSC complex does is about how much Rheb-GTP there is.

What does it mean that TSC2 acts as a GAP for Rheb?
  1. It binds Rheb and physically blocks its interaction with mTORC1
  2. It accelerates Rheb's hydrolysis of its own bound GTP, pushing Rheb toward the GDP state
  3. It loads Rheb with fresh GTP
  4. It degrades Rheb protein

Answer: B. A GTPase-activating protein speeds up a reaction the GTPase already performs. That is why the TSC complex sets a rate rather than flipping a switch, and why the amount of Rheb-GTP is a balance.

What holds the Rag GTPases at the lysosomal surface?
  1. Rheb
  2. The Ragulator complex
  3. RAPTOR
  4. TFEB

Answer: B. The Rags are not membrane-anchored on their own. Ragulator tethers them to the lysosome and acts on their nucleotide state, which is what makes the surface a platform rather than a passive membrane.

Which amino acid does Sestrin2 bind?
  1. Arginine
  2. Leucine
  3. Glutamine
  4. Methionine

Answer: B. Sestrin2 is the leucine-binding protein of this set; CASTOR1 handles arginine and SAMTOR reads S-adenosylmethionine, the metabolite downstream of methionine.

‘‘mTOR controls growth’’ — which of these does the evidence behind this claim not support?
  1. That several unrelated kinds of information reach the same kinase, which is what makes it worth calling an integrator
  2. That mTOR activity shifts the balance between building programs and recycling programs in the systems where this has been measured
  3. That mTOR is the only route to growth, or that removing it stops growth in every cell

Answer: C. Almost everything in this lesson was worked out in cultured cells and in laboratory animals, and the summary sentence compresses a network with many parallel inputs and outputs into one verb. The compression is useful for learning and it is not a finding; when you meet a claim about mTOR and human health later in the Atlas, the evidence tier attached to it is doing real work.

‘What ‘mTORC1-selective’ can and cannot mean’ — which of these does the evidence behind this claim not support?
  1. That the two complexes differ in composition and in substrates, and that the difference is visible both chemically and genetically
  2. That rapamycin blocks some mTORC1 outputs much more completely than others
  3. That any compound is selective as a property independent of dose, duration and cell type

Answer: C. Every study cited in this lesson sits at tier D or C: biochemistry, cell work and mouse genetics. That is the honest state of complex-level evidence. Selectivity in particular is a claim about an experiment, and the experiment has a duration — which is exactly the thing summaries drop.

‘What this lesson's evidence supports, and what it does not’ — which of these does the evidence behind this claim not support?
  1. The TSC complex acts as a GAP for Rheb, which is how growth-factor and stress signals reach the pathway
  2. Rheb contributes to mTORC1 activation, and its GTP-bound state is the form that does it
  3. Localisation is part of the mechanism rather than a detail drawn on top of it
  4. That Rheb is the only determinant of mTORC1 activity

Answer: D. Several of the foundational experiments in this lesson raised the amount of a protein far above its normal level in order to see its effect. That design is informative about what a protein can do and much weaker evidence about what it normally does — and the studies below sit at tier D and C, which marks the design, not the quality.

Which limitation is the one this study states about itself? — Read the design, not the wording.

mTOR regulates memory CD8 T-cell differentiation (2009, Mouse)
mTOR is a key regulator of memory CD8 T-cell differentiation; rapamycin enhances memory responses.
Sample: not stated

  1. mTOR knockdown does not completely mimic rapamycin treatment.
  2. No lifespan or healthspan endpoint. Side-effect endpoints only; benefit retention is inferred, not tested.
  3. FDA changed the primary endpoint between phase 2b and phase 3 (symptom-based vs lab-confirmed), complicating comparison; positive biomarker did not translate to clinical benefit.
  4. Inference from a single exceptional responder; basis of mTOR-inhibitor sensitivity incompletely understood; short drug exposure in some.

Answer: A. Its own limitation, as recorded in the Atlas: mTOR knockdown does not completely mimic rapamycin treatment.

Which limitation is the one this study states about itself? — Read the design, not the wording.

Alternative rapamycin treatment regimens mitigate the impact of rapamycin on glucose homeostasis and the immune system (2015, Mouse)
Intermittent rapamycin regimens (weekly, or every 5 days) largely spared glucose tolerance, pyruvate tolerance, fasting glucose and insulin, beta-cell function and the immune system, while still inhibiting mTORC1 -- unlike daily dosing, which impaired all of…
Sample: 9-11 male C57BL/6J mice per treatment group for glucose tolerance tests; 3-6 mice per group for blood rapamycin concentration; 4-9 mice per group for…

  1. Histological evaluation of cataracts was limited to a small number of specimens; site-to-site variation was considerable for spontaneous activity; some observed trends did not reach statistical significance.
  2. Specific inhibitors of mTORC2 are unavailable, precluding pharmacological testing. Differences in genetic background between Rictor +/- mice and mtor/Raptor/mlst8 heterozygous mice precluded prostate cancer…
  3. Inference from rare responders; basis of mTOR-inhibitor sensitivity beyond TSC1/2 loss still incompletely understood.
  4. No lifespan or healthspan endpoint. Side-effect endpoints only; benefit retention is inferred, not tested.

Answer: D. Its own limitation, as recorded in the Atlas: No lifespan or healthspan endpoint. Side-effect endpoints only; benefit retention is inferred, not tested.

How settled is this, as the field stands? — 4E-BP1 inhibits eIF4E

Unphosphorylated 4E-BP1 clamps onto eIF4E and stops it assembling the cap-binding complex. mTORC1 phosphorylates 4E-BP1 to release that grip - so the brake on translation is released, not the accelerator pressed.

  1. Established
  2. Emerging
  3. Contested
  4. Open — no answer either way

Answer: A. Curated in the Atlas as consensus established, mechanistic evidence high, human relevance plausible.

How settled is this, as the field stands? — 4E-BP1 activates Longevity

Keeping 4E-BP active extends lifespan under dietary restriction in flies, by preserving mitochondrial activity. One of the few places where a single downstream node, not the whole pathway, carries the lifespan effect.

  1. Established
  2. Emerging
  3. Contested
  4. Open — no answer either way

Answer: B. Curated in the Atlas as consensus emerging, mechanistic evidence medium, human relevance untested. Scope: Fly only, and only under dietary restriction - the effect was not seen on a full diet.

Which study in the Atlas is this claim standing on? — 4E-BP1 inhibits eIF4E

Unphosphorylated 4E-BP1 clamps onto eIF4E and stops it assembling the cap-binding complex. mTORC1 phosphorylates 4E-BP1 to release that grip - so the brake on translation is released, not the accelerator pressed.

  1. HOS2009 — Nutrient-dependent mTORC1 association with the ULK1-Atg13-FIP200 complex required for… (2009)
  2. ROM2001 — Mediation of IGF-1-induced skeletal myotube hypertrophy by PI(3)K/Akt/mTOR and… (2001)
  3. MAX2009 — Molecular mechanisms of mTOR-mediated translational control (2009)
  4. ZID2009 — 4E-BP extends lifespan upon dietary restriction by enhancing mitochondrial activity in… (2009)

Answer: C. The Atlas records MAX2009, THO2012, SCH2003 as evidence for this step.

4E-BP1 → eIF4E
  1. activates
  2. inhibits

Answer: B. Unphosphorylated 4E-BP1 clamps onto eIF4E and stops it assembling the cap-binding complex. mTORC1 phosphorylates 4E-BP1 to release that grip - so the brake on translation is released, not the accelerator pressed.

4E-BP1 → Longevity
  1. activates
  2. inhibits

Answer: A. Keeping 4E-BP active extends lifespan under dietary restriction in flies, by preserving mitochondrial activity. One of the few places where a single downstream node, not the whole pathway, carries the lifespan effect.

Akt/PKB → TSC1/TSC2
  1. activates
  2. inhibits

Answer: B. Akt phosphorylates TSC2, disabling the complex. Growth-factor signalling works by removing a brake, not by pressing an accelerator.

AMPK → Mitophagy
  1. activates
  2. inhibits

Answer: A. The LKB1-AMPK axis keeps damaged mitochondria being cleared; lose it and mitochondrial DNA leaks into the cytosol and inflames the cell.

Routes to rebuild

In the game these arrive as loose parts to be put back in order. Here they are written out, with the answer.

How does a cell know it has enough raw material to grow?

Leucine ⊣ Sestrin2 ⊣ GATOR2 ⊣ GATOR1 ⊣ Rag GTPases

How does a cell learn that it is allowed to grow?

Growth hormone / IGF-1 axis → PI3K → Akt/PKB ⊣ TSC1/TSC2 ⊣ Rheb → mTORC1 → S6K1 ⊣ IRS-1 / IRS-2 — 2 of these steps is indirect: it runs through a step this route does not draw.